NNAT (neuronatin) has long (alpha) and short (beta) transcripts. Both were monoallelically expressed
in three fetuses and one dam in a wide variety of tissues (lung, brain, liver, kidney, muscle,
ovary, pituitary, mammary gland,
eye, intestine, endometrium, lung, spleen, heart). The expressed allele was said to be paternal but
this was not
obviously demonstrated (Zaitoun
& Khatib 2006).
NNAT expression was detected in embryos arising from in-vitro fertilisation, but not in those from
parthenogenetic activation, consistent with the imprinting (paternal expression) of NNAT in bovine
NT et al, 2004).