Tssc4 showed a significant bias toward maternal expression in the placenta (Paulsen
M et al, 2000; Umlauf
D et al, 2004).
Biased maternal expression was also detected in the visceral yolk sac (Hudson QJ et al, 2011).
It is thought not to be imprinted in humans.
Okae H et al, (2012) confirmed
imprinted maternal expression of this gene in placenta and trophoblast stem cells (TS).