Ccdc40 (coiled-coil domain containing 40) showed biased paternal expression in E15 brain, E17 brain
( DeVeale B, 2012 ). Ccdc40 was one of six
validated imprinted genes among 53 candidate imprinted genes detected in brain transcriptomics
analysis by DeVeale B et al.
Ccdc40 was one of 1308 candidate imprinting loci described by
Gregg C et al (2010).